/**
 * Biology Hub
 *
 * Three-column biology and animal-vocabulary entry page.
 */

body.biology-hub-page {
    --bio-line: #ddd2c3;
    --bio-muted: #76675b;
    --bio-orange: #d97816;
    --bio-brown: #8b6244;
}

body.biology-hub-page * { box-sizing: border-box; }

body.biology-hub-page main.biology-hub-shell {
    display: grid;
    width: 100%;
    max-width: none;
    margin: 0;
    grid-template-columns: 250px minmax(0, 1fr) 300px;
    align-items: stretch;
    border: 1px solid var(--bio-line);
    background: #fff;
}

body.biology-hub-page .biology-hub-left,
body.biology-hub-page .biology-hub-right {
    padding: 18px 18px 28px;
    background: #fbf9f5;
}

body.biology-hub-page .biology-hub-left { border-right: 1px solid var(--bio-line); }
body.biology-hub-page .biology-hub-right { border-left: 1px solid var(--bio-line); }
body.biology-hub-page .biology-hub-main { min-width: 0; padding: 18px 22px 38px; }
body.biology-hub-page .hub-hero { margin: 0 0 18px; padding: clamp(2rem, 5vw, 3.4rem) 1.25rem clamp(2.15rem, 4vw, 3rem); }

body.biology-hub-page .biology-panel-title {
    margin: 0 0 10px;
    padding-bottom: 8px;
    border-bottom: 1px solid var(--bio-line);
    color: var(--bio-orange);
    font-size: 1rem;
    font-weight: 800;
}

body.biology-hub-page .biology-side-section { margin-top: 26px; }
body.biology-hub-page .biology-side-list { display: grid; gap: 8px; }

body.biology-hub-page .biology-side-link,
body.biology-hub-page .biology-info-card {
    display: block;
    padding: 11px 12px;
    border: 1px solid var(--bio-line);
    border-radius: 9px;
    background: #fff;
    color: inherit;
    text-decoration: none;
}

body.biology-hub-page a.biology-side-link:hover,
body.biology-hub-page a.biology-side-link:focus-visible,
body.biology-hub-page a.biology-info-card:hover,
body.biology-hub-page a.biology-info-card:focus-visible {
    border-color: var(--bio-orange);
    box-shadow: 0 4px 12px rgba(70, 48, 30, 0.08);
    color: inherit;
    text-decoration: none;
}

body.biology-hub-page .biology-side-link strong,
body.biology-hub-page .biology-info-card strong { display: block; color: #49372b; }

body.biology-hub-page .biology-side-link small,
body.biology-hub-page .biology-info-card small {
    display: block;
    margin-top: 3px;
    color: var(--bio-muted);
    font-size: 0.76rem;
    line-height: 1.45;
}

body.biology-hub-page .biology-featured {
    margin: 0;
    padding: 17px;
    border: 1px solid #d7c2a8;
    border-radius: 14px;
    background: #fffaf1;
}

body.biology-hub-page .biology-featured__header { display: flex; gap: 16px; margin-bottom: 13px; }
body.biology-hub-page .biology-featured__header h2,
body.biology-hub-page .biology-section__header h2 { margin: 0; color: var(--bio-orange); font-size: 1.25rem; }

body.biology-hub-page .biology-featured__header p,
body.biology-hub-page .biology-section__header p {
    margin: 5px 0 0;
    color: var(--bio-muted);
    font-size: 0.86rem;
    line-height: 1.65;
}

body.biology-hub-page .biology-featured__grid {
    display: grid;
    grid-template-columns: repeat(3, minmax(0, 1fr));
    gap: 10px;
}

body.biology-hub-page .biology-featured-card {
    display: block;
    min-width: 0;
    padding: 13px;
    border: 1px solid var(--bio-line);
    border-left: 5px solid var(--collection-accent);
    border-radius: 10px;
    background: #fff;
    color: inherit;
    text-decoration: none;
}

body.biology-hub-page .biology-featured-card:hover,
body.biology-hub-page .biology-featured-card:focus-visible {
    border-color: var(--collection-accent);
    box-shadow: 0 5px 14px rgba(70, 48, 30, 0.09);
    color: inherit;
    text-decoration: none;
}

body.biology-hub-page .biology-featured-card strong { display: block; color: #3f3026; line-height: 1.35; }
body.biology-hub-page .biology-featured-card span { display: block; margin-top: 5px; color: var(--bio-muted); font-size: 0.76rem; line-height: 1.45; }

body.biology-hub-page .biology-section { margin-top: 30px; }
body.biology-hub-page .biology-section__header { margin-bottom: 13px; padding-bottom: 9px; border-bottom: 1px solid var(--bio-line); }

body.biology-hub-page .biology-glossary-grid,
body.biology-hub-page .biology-animal-grid {
    display: grid;
    grid-template-columns: repeat(2, minmax(0, 1fr));
    gap: 12px;
}

body.biology-hub-page .biology-glossary-card,
body.biology-hub-page .biology-animal-card {
    min-width: 0;
    padding: 16px;
    border: 1px solid var(--bio-line);
    border-radius: 12px;
    color: inherit;
}

body.biology-hub-page .biology-glossary-card {
    display: block;
    border-left: 5px solid var(--bio-brown);
    background: #fff;
    text-decoration: none;
}

body.biology-hub-page .biology-glossary-card:hover,
body.biology-hub-page .biology-glossary-card:focus-visible {
    border-color: var(--bio-brown);
    box-shadow: 0 5px 14px rgba(70, 48, 30, 0.09);
    color: inherit;
    text-decoration: none;
}

body.biology-hub-page .biology-animal-card {
    overflow: hidden;
    border-top: 6px solid var(--animal-accent);
    background: var(--animal-bg);
}

body.biology-hub-page .biology-glossary-card__meta,
body.biology-hub-page .biology-animal-card__meta {
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: 12px;
    margin-bottom: 8px;
}

body.biology-hub-page .biology-badge {
    display: inline-block;
    padding: 3px 7px;
    border: 0;
    border-radius: 4px;
    background: #eee8e2;
    color: var(--animal-accent, var(--bio-brown));
    font-size: 0.7rem;
    font-weight: 800;
}

body.biology-hub-page .biology-count {
    color: var(--animal-accent, var(--bio-brown));
    font-size: 0.77rem;
    font-weight: 800;
    text-align: right;
}

body.biology-hub-page .biology-glossary-card h3,
body.biology-hub-page .biology-animal-card h3 { margin: 0; color: #3c2d23; font-size: 1.08rem; line-height: 1.35; }

body.biology-hub-page .biology-glossary-card p,
body.biology-hub-page .biology-animal-card p {
    margin: 7px 0 0;
    color: var(--bio-muted);
    font-size: 0.87rem;
    line-height: 1.6;
}

body.biology-hub-page .biology-page-links { display: flex; flex-wrap: wrap; gap: 7px; margin-top: 12px; }

body.biology-hub-page .biology-page-link {
    display: inline-flex;
    align-items: center;
    min-height: 30px;
    padding: 6px 10px;
    border: 1px solid #ddcfc2;
    border-radius: 999px;
    background: #fff;
    color: #56463a;
    font-size: 0.77rem;
    box-shadow:
        inset 0 1px 0 rgba(255, 255, 255, 0.85),
        0 1px 2px rgba(60, 45, 35, 0.12),
        0 2px 5px rgba(60, 45, 35, 0.08);
    font-weight: 700;
    line-height: 1.35;
    text-decoration: none;
}

body.biology-hub-page .biology-page-link:hover,
body.biology-hub-page .biology-page-link:focus-visible {
    border-color: var(--animal-accent, var(--bio-brown));
    background: var(--animal-accent, var(--bio-brown));
    color: #fff;
    box-shadow:
        inset 0 1px 0 rgba(255, 255, 255, 0.22),
        0 2px 4px rgba(60, 45, 35, 0.2),
        0 4px 9px rgba(60, 45, 35, 0.12);
    text-decoration: none;
}

body.biology-hub-page .biology-card-action {
    display: inline-block;
    margin-top: 13px;
    color: var(--animal-accent, var(--bio-brown));
    font-size: 0.82rem;
    font-weight: 800;
    text-decoration: none;
}

body.biology-hub-page a.biology-card-action:hover,
body.biology-hub-page a.biology-card-action:focus-visible { color: var(--bio-orange); text-decoration: underline; }

/* Shared topic-hub sidebar contract: destination rows, compact terms, guidance, and related rows. */
body.biology-hub-page .biology-side-section { margin-top: 1.55rem; }
body.biology-hub-page .biology-side-list { gap: 0.12rem; }

body.biology-hub-page .biology-side-link--destination {
    padding: 8px 4px 8px 11px;
    border: 0;
    border-left: 3px solid transparent;
    border-radius: 0;
    background: transparent;
}

body.biology-hub-page .biology-side-link--destination:hover,
body.biology-hub-page .biology-side-link--destination:focus-visible,
body.biology-hub-page .biology-side-link--destination[aria-current="page"] {
    border-left-color: var(--bio-brown);
    background: #f6ede4;
    box-shadow: none;
}

body.biology-hub-page .biology-info-card--term {
    margin: 3px 0;
    padding: 8px 10px;
    border-left: 3px solid var(--bio-brown);
    border-radius: 7px;
}

body.biology-hub-page .biology-info-card--guidance {
    padding: 0 0 12px;
    border: 0;
    border-bottom: 1px solid var(--bio-line);
    border-radius: 0;
    background: transparent;
}

body.biology-hub-page .biology-info-card--guidance + .biology-info-card--guidance { padding-top: 12px; }

body.biology-hub-page .biology-side-link--related {
    display: grid;
    grid-template-columns: minmax(0, 1fr) auto;
    align-items: center;
    justify-content: space-between;
    gap: 9px;
    padding: 10px 2px;
    border: 0;
    border-bottom: 1px solid #e3d8cc;
    border-radius: 0;
    background: transparent;
}

body.biology-hub-page .biology-side-link--related::after {
    grid-row: 1 / span 2;
    grid-column: 2;
    flex: 0 0 auto;
    color: var(--bio-brown);
    content: "\2192";
    font-size: 1rem;
}

body.biology-hub-page .biology-side-link--related:hover,
body.biology-hub-page .biology-side-link--related:focus-visible {
    border-bottom-color: #bfa178;
    background: transparent;
    box-shadow: none;
    color: var(--bio-brown);
}

body.biology-hub-page .biology-side-link--destination:focus-visible,
body.biology-hub-page .biology-side-link--related:focus-visible,
body.biology-hub-page .biology-info-card--term:focus-visible {
    outline: 2px solid rgba(217, 120, 22, 0.5);
    outline-offset: 2px;
}

body.biology-hub-page .biology-animal-intro {
    padding: 18px;
    border: 1px solid #d4c0a9;
    border-radius: 14px;
    background: #fffaf3;
}

body.biology-hub-page .biology-plant-intro {
    border-color: #bfd0b8;
    background: #f8fcf6;
}

body.biology-hub-page .biology-plant-intro h2 { color: #41683c; }

body.biology-hub-page .biology-plant-intro .biology-animal-intro__stats {
    color: #41683c;
}

body.biology-hub-page .biology-animal-intro__top { display: flex; align-items: flex-start; justify-content: space-between; gap: 18px; }
body.biology-hub-page .biology-animal-intro h2 { margin: 0; color: #68452f; font-size: 1.45rem; }
body.biology-hub-page .biology-animal-intro p { margin: 7px 0 0; color: var(--bio-muted); line-height: 1.65; }

body.biology-hub-page .biology-animal-intro__stats {
    flex: 0 0 auto;
    padding: 4px 0;
    border: 0;
    border-bottom: 1px solid currentColor;
    border-radius: 0;
    background: transparent;
    color: #6e4f36;
    font-size: 0.82rem;
    font-weight: 800;
    text-decoration: none;
    text-underline-offset: 0.18em;
}

body.biology-hub-page .biology-animal-intro__stats:hover,
body.biology-hub-page .biology-animal-intro__stats:focus-visible {
    color: var(--bio-orange);
    text-decoration: underline;
}

/* Biology Hub domain overviews: detailed route cards live on the child hubs. */
body.biology-hub-page .biology-domain-card {
    --domain-accent: #765038;
    --domain-ink: #513827;
    --domain-line: #d8c2ad;
    --domain-soft: #fff9f2;
    overflow: hidden;
    padding: 20px;
    border: 1px solid var(--domain-line);
    border-radius: 14px;
    background: var(--domain-soft);
}

body.biology-hub-page .biology-domain-card--plant {
    --domain-accent: #477a4a;
    --domain-ink: #355d38;
    --domain-line: #bfd0b8;
    --domain-soft: #f8fcf6;
}

body.biology-hub-page .biology-domain-card__head h2 {
    margin: 0;
    color: var(--domain-ink);
    font-size: 1.45rem;
    line-height: 1.35;
}

body.biology-hub-page .biology-domain-card__head p {
    max-width: 720px;
    margin: 7px 0 0;
    color: var(--bio-muted);
    font-size: 0.88rem;
    line-height: 1.65;
}

body.biology-hub-page .biology-domain-routes {
    display: grid;
    grid-template-columns: repeat(2, minmax(0, 1fr));
    gap: 10px;
    margin-top: 16px;
}

/* Each route carries the accent its card uses on the child hub, so the colour
   is a wayfinding cue rather than decoration. */
body.biology-hub-page .biology-domain-route {
    display: grid;
    grid-template-columns: auto minmax(0, 1fr);
    align-items: start;
    gap: 10px;
    min-width: 0;
    padding: 12px;
    border: 1px solid var(--domain-line);
    border-left: 4px solid var(--route-accent, var(--domain-accent));
    border-radius: 9px;
    background: rgba(255, 255, 255, 0.72);
}

body.biology-hub-page .biology-domain-route__number {
    min-width: 32px;
    padding: 3px 6px;
    border-radius: 4px;
    background: #eee8e2;
    color: var(--route-accent, var(--domain-accent));
    font-size: 0.7rem;
    font-weight: 850;
    line-height: 1.35;
    text-align: center;
}

body.biology-hub-page .biology-domain-route h3 {
    margin: 0;
    color: #3c2d23;
    font-size: 0.92rem;
    line-height: 1.4;
}

body.biology-hub-page .biology-domain-route h3 a {
    color: var(--domain-ink);
    text-decoration: underline;
    text-decoration-color: transparent;
    text-underline-offset: 0.18em;
}

body.biology-hub-page .biology-domain-route h3 a:hover,
body.biology-hub-page .biology-domain-route h3 a:focus-visible {
    color: var(--domain-accent);
    text-decoration-color: currentColor;
}

body.biology-hub-page .biology-domain-route p {
    margin: 4px 0 0;
    color: var(--bio-muted);
    font-size: 0.76rem;
    line-height: 1.5;
}

/* Direct glossary links are the primary action on this hub: one click to terms. */
body.biology-hub-page .biology-domain-route__links {
    display: flex;
    flex-wrap: wrap;
    gap: 6px;
    margin: 10px 0 0;
    padding: 0;
    list-style: none;
}

body.biology-hub-page .biology-domain-route__links a {
    display: inline-flex;
    align-items: center;
    min-height: 30px;
    padding: 6px 10px;
    border: 1px solid #ddcfc2;
    border-radius: 999px;
    background: #fff;
    color: #56463a;
    font-size: 0.79rem;
    box-shadow:
        inset 0 1px 0 rgba(255, 255, 255, 0.85),
        0 1px 2px rgba(60, 45, 35, 0.12);
    font-weight: 700;
    line-height: 1.35;
    text-decoration: none;
}

body.biology-hub-page .biology-domain-route__links a:hover,
body.biology-hub-page .biology-domain-route__links a:focus-visible {
    border-color: var(--domain-accent);
    background: var(--domain-accent);
    color: #fff;
    box-shadow:
        inset 0 1px 0 rgba(255, 255, 255, 0.22),
        0 2px 4px rgba(60, 45, 35, 0.2);
    text-decoration: none;
}

body.biology-hub-page .biology-domain-route h3 a:focus-visible,
body.biology-hub-page .biology-domain-route__links a:focus-visible {
    outline: 2px solid rgba(217, 120, 22, 0.5);
    outline-offset: 2px;
}

body.biology-hub-page .biology-domain-card__footer {
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: 18px;
    margin-top: 16px;
    padding-top: 14px;
    border-top: 1px solid var(--domain-line);
}

body.biology-hub-page .biology-domain-card__footer p {
    max-width: 560px;
    margin: 0;
    color: var(--bio-muted);
    font-size: 0.8rem;
    line-height: 1.55;
}

/* Secondary route: the child hub is for the full map and index, not a gate. */
body.biology-hub-page .biology-domain-action {
    flex: 0 0 auto;
    padding: 4px 0;
    border: 0;
    border-bottom: 1px solid currentColor;
    border-radius: 0;
    background: transparent;
    box-shadow: none;
    color: var(--domain-accent);
    font-size: 0.8rem;
    font-weight: 800;
    line-height: 1.35;
    text-decoration: none;
    text-underline-offset: 0.18em;
}

body.biology-hub-page .biology-domain-action:hover,
body.biology-hub-page .biology-domain-action:focus-visible {
    color: var(--bio-orange);
    text-decoration: none;
}

body.biology-hub-page .biology-domain-action:focus-visible {
    outline: 2px solid rgba(217, 120, 22, 0.55);
    outline-offset: 3px;
}

body.biology-hub-page .biology-animal-grid { margin-top: 14px; }
body.biology-hub-page .biology-search { display: grid; grid-template-columns: minmax(0, 1fr) auto; gap: 8px; }
body.biology-hub-page .biology-search input { width: 100%; min-width: 0; padding: 10px 11px; border: 1px solid #cbbba9; border-radius: 8px; font: inherit; }
body.biology-hub-page .biology-search input:focus { outline: 2px solid rgba(217, 120, 22, 0.35); outline-offset: 2px; }

body.biology-hub-page .biology-search button {
    padding: 10px 15px;
    border: 1px solid #bfa178;
    border-radius: 8px;
    background: #9a7b5c;
    color: #fff;
    font: inherit;
    font-weight: 800;
}

body.biology-hub-page .biology-search button:hover,
body.biology-hub-page .biology-search button:focus-visible { background: #7f5f43; }

body.biology-hub-page .biology-search-status { margin: 9px 0 0; color: #9d3b25; font-size: 0.82rem; line-height: 1.5; }
body.biology-hub-page .is-search-hidden { display: none; }

body.biology-hub-page .biology-featured-card:focus-visible,
body.biology-hub-page .biology-glossary-card:focus-visible,
body.biology-hub-page .biology-page-link:focus-visible,
body.biology-hub-page .biology-card-action:focus-visible,
body.biology-hub-page .biology-animal-intro__stats:focus-visible {
    outline: 2px solid rgba(217, 120, 22, 0.55);
    outline-offset: 3px;
}

@media (max-width: 1180px) {
    body.biology-hub-page main.biology-hub-shell { grid-template-columns: 250px minmax(0, 1fr); }
    body.biology-hub-page .biology-hub-right { grid-column: 1 / -1; border-top: 1px solid var(--bio-line); border-left: 0; }
}

@media (max-width: 900px) {
    body.biology-hub-page main.biology-hub-shell { grid-template-columns: 1fr; }
    body.biology-hub-page .biology-hub-main { order: 1; }
    body.biology-hub-page .biology-hub-left { order: 2; border-top: 1px solid var(--bio-line); border-right: 0; }
    body.biology-hub-page .biology-hub-right { order: 3; }
    body.biology-hub-page .biology-featured__grid,
    body.biology-hub-page .biology-glossary-grid,
    body.biology-hub-page .biology-animal-grid { grid-template-columns: 1fr; }
    body.biology-hub-page .biology-animal-intro__top { display: block; }
    body.biology-hub-page .biology-animal-intro__stats { display: inline-block; margin-top: 12px; }
    body.biology-hub-page .biology-domain-card__footer { align-items: flex-start; flex-direction: column; }
}

@media (max-width: 560px) {
    body.biology-hub-page .biology-hub-left,
    body.biology-hub-page .biology-hub-right,
    body.biology-hub-page .biology-hub-main { padding-right: 14px; padding-left: 14px; }
    body.biology-hub-page .biology-featured,
    body.biology-hub-page .biology-animal-intro,
    body.biology-hub-page .biology-glossary-card,
    body.biology-hub-page .biology-animal-card { padding: 13px; }
    body.biology-hub-page .biology-glossary-card__meta,
    body.biology-hub-page .biology-animal-card__meta { align-items: flex-start; flex-direction: column; gap: 6px; }
    body.biology-hub-page .biology-count { text-align: left; }
    body.biology-hub-page .biology-domain-card { padding: 15px; }
    body.biology-hub-page .biology-domain-routes { grid-template-columns: 1fr; }
}

/* Multidimensional sidebar treatment from the Biology Hub reference. */
body.biology-hub-page .bio-sidebar-section + .bio-sidebar-section { margin-top: 1.55rem; }
body.biology-hub-page .bio-overview-link { display: block; margin: 0 0 1rem; padding: 0.42rem 0; border-bottom: 1px solid var(--bio-line); color: #49372b; font-size: 0.88rem; font-weight: 750; text-decoration: none; }
body.biology-hub-page .bio-overview-link::before { margin-right: 0.38rem; color: var(--bio-orange); content: "\2302"; }
body.biology-hub-page .bio-overview-link[aria-current="page"] { color: var(--bio-orange); }
body.biology-hub-page .bio-toc { display: grid; gap: 1.15rem; }
body.biology-hub-page .bio-toc-section { min-width: 0; }
body.biology-hub-page .bio-toc-section__title { margin: 0 0 0.32rem; padding: 0 0 0.38rem; border-bottom: 1px solid var(--bio-line); color: var(--dimension-accent, var(--bio-orange)); font-size: 0.84rem; font-weight: 850; line-height: 1.35; }
body.biology-hub-page .bio-toc-list { display: grid; gap: 0.12rem; margin: 0; padding: 0; list-style: none; }
body.biology-hub-page .bio-toc-link { display: block; padding: 0.34rem 0; color: #4f4339; font-size: 0.86rem; line-height: 1.45; text-decoration: none; }
body.biology-hub-page .bio-featured-stack { display: grid; gap: 0.58rem; }
body.biology-hub-page .bio-featured-link { display: block; padding: 0.7rem 0.72rem; border: 1px solid var(--bio-line); border-left: 3px solid var(--bio-orange); border-radius: 6px; background: #fff; color: inherit; text-decoration: none; }
body.biology-hub-page .bio-featured-link strong { display: block; color: #49372b; font-size: 0.86rem; line-height: 1.4; }
body.biology-hub-page .bio-featured-link small { display: block; margin-top: 0.16rem; color: var(--bio-muted); font-size: 0.73rem; line-height: 1.42; }
body.biology-hub-page .bio-overview-link:hover, body.biology-hub-page .bio-overview-link:focus-visible, body.biology-hub-page .bio-toc-link:hover, body.biology-hub-page .bio-toc-link:focus-visible { color: var(--bio-orange); text-decoration: underline; text-underline-offset: 0.18em; }
body.biology-hub-page .bio-overview-link:focus-visible, body.biology-hub-page .bio-toc-link:focus-visible, body.biology-hub-page .bio-featured-link:focus-visible { outline: 2px solid rgba(114, 84, 61, 0.28); outline-offset: 2px; }